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. ( ) 03.02.06 - - 2012 л - ( л - ) :, :,,, л - :,,, л :...,. л28 2012. 10-00 307.012.01 л - : 690091,.,, 4. (423)2300-751. E-mail: dissovet@tinro.ru л - л25 2012
....,,,,, (, 2006). 1980-., - (., 2010)
-., XIX (, 1998;., 2004).,,.,,,.,
.. ( )
:
1.
2.
3.
4. -
.. Bathyraja parmifera, Alcichthys elongatus Enophrys diceraus, Gymnocanthus pistilliger, G. herzensteini G. detrisus, Icelus cataphractus, Myoxocephalus brandti, M. polyacanthocephalus - M. jaok, - Hemitripterus villosus, Liparis ochotensis, Bothrocara hollandi, Liopsetta pinnifasciata, Platichthys stellatus, Limanda punctatissima, Pseudopleuronectes yokohamae Acanthopsetta nadeshnyi, Stichaeus grigorjewi
Clupea pallasii, Tribolodon brandti, Eleginus gracilis, Gadus macrocephalus, Theragra chalcogramma, Hexagrammos stelleri Pleurogrammus azonus, Lumpenus sagitta, Ps. herzensteini, L. aspera, Glyptocephalus stelleri, Cleisthenes herzensteini Hippoglossoides dubius
, ( ).. -,
....,.,
,.,, -. -
. 1. 32 11, 9 - 12 Ц. (34.4%), (58.5%) II-III.
2.. ( 32%),,, - (18.3%) (17%).
3. -.,,
., ( ) - 2002-2009.,.,, 1994-19
. л (, 1996); PICES (,, 2000); л (, 2001); - л., (, 2008; -, 2011); л (, 2010); - (2009, 2010, 2011)
. 19, 13 -,.
.,, (274, 41 ). 200, 44 31
......,....
,..,....,....,....,..,..............,.....,..,.
......,,,.....,,...........,..........
1.,. (5 - 600 ) ( - ) -5005 (2002-2007.), л (2007.) л (2009.) (. 1)
23. 27.1- / 10
. 1. - : 1 Ц., 2 Ц, 3 Ц, 4 Ц.
( Е, 1974; Е, 1986). ( ) 1,.. - 10%.. 0.01. ( % ). 2002-2009. 6481 32 (
1)., 1994-1999.. 1996-2005.
,. 2002-2009
32 1 180 79 327 1 382 1 552 1 156 426 1 120 2 110 3 110 1 163 2 266 3 145 2 200 3 - 2 156 64 ( ).
(1949),.... (1999):,.. (1986):
D = 16*W0,18*ym0,6*, (1) Dx - 90% ( ), W - ( ), ym - ( ) ( ), Ц
(Jones, 1974):
= exp [0,0806*(20 - Q)], (2) Q Ц,
: ( 50 ), (50-200 ) ( 200 )
( ).. (1952) ( ). 33%, 34 66% -, 66% -
.. Ц.. (1957).. (2002):
a n = *D* 1* 2, (3) a m a n Ц, - a m ( ), D Ц.. (1952) ( 1), 1 2 - ( % ). 1000, 1000 5000 Ц, 5000 Ц
(Cortes, 1999):
n TLk = 1 + ( Pj*TLj), (4) j= TLk - k, PJ - j k, n Ц, TLj - j
, ( I ) -, - ( I-II ) Ц, ( II, I ) - (, 1980;, 2002;, 2006.).,, (4) ( - )
2. 2.1.... ( -, -, - ), ( -, - ) (, 1983)., (,,,.), Ц. ( 1860.). 312. 20-40-. (233) (236) (., 2009)
,,., ( ), - (,,.), - ( Sardinops melanostictus), ( ) (, 1998;., 2004)
., 5-50. ( 200-600 ) (, 2004, 2005, 2006)
2001-2009. 147, 36., 78.5., (28.5%), (22.8%), (18.7%) (14.8%), (17.1%), (14.4%), (8.3%) - (8.3%). 10.3 / 2, - 5.7 / 2, - 3.5 / 2.,, (23.4%), (19%), (9.6%), (8.8%) (8.4%) (,, 2012)
2.2.... (, 2010), ( 2008 / 3), (40.2%), (32.8%) (19%),,. ( 849 / 3 ), (51.6%), (8.7%), (11.4%), (24.2%) ( 75%),. 1.
л 265.4 /., 1000 / 2. 7-280 1.., - 1.6., (30.7%), (24.4%), (16.9%) (8.3%) (., 2005).. 1930-. (,, 1941), (200-600 ) 87 / 2, ( ) - 34.5 / 2
,.,,, ( ),, - (, 2001;, 2002). ( 699.4 501 / ) (., 2008)
,,,,.., - - Chionoecetes opilio - 12 600, 100-350 (, 2008;, 2011). - - Sclerocrangon salebrosa - 30 110, (, 2008). Pandalus borealis P. hypsinotus, (,, 2010). - Todarodes pacificus (, 2006)
3.. 32,.
(57.4%) (28.9%), - (10.7%). 21-40 ( ), (41-80 ) -, -. 81-1 ( ), 101-1 - ( ). 2 0.3% 0.9 %
(32.1%), (24.3%) (19.9%), - (16.6%) (5.3%). 11-15. 16-25, (26-30 ). 4.5% 7.9 2.8%
( 27.3 27.2%), - (17%), (16.7%) (11.8%). 16-20, (21-30 ), 30 -. ( 6.8 0.7%) 2.9%
8-35 (33.3%), (25.2%) (21.2%).,,. 8-15,,, ( ). 5.3%, 6.4 2.6%
(60.5%),,,,. (20.6%), -, (11.9%),. 21-30,,,,. (31-70 ),. 70. 1.5-3.2% ( 2.7%)
(40.1%) (20.1%), - (11.3%), (7.8%), (5.6%), (6.8%) (4.6%). 11-20, Ц,.,,,. 60. 3% 4.1 2.1 %
(46.4%) (29.4%). 11-20,,, (21-40 ) - ( ), -,,. 11.2 5.7% ( 7.2%)
(23.1%), (22.9%) (20.4%), - (11.6%),, (10.1%) (5.7%). 21-30,, (,,, -, ). 41-50. 4.1-5.6% ( 4.3%)
( - ) - 69.2% ( ) - 20.5%. 11-20,. - 31-40, - ( 40 ). 4.1 2.8% 3%
(43.7%) (34%). (6.7%), (4.9%), (4.1%) (3.3%). 16-20, -. 21-25, (26-30 ) - -. 1%, 0.7-1.4%
(54.4%), - (17.1%), (8.2%), (11.5%) (5.3%). 11-15, ( ). (21-25 ). (3.5-3.3%)
(37.2%), Engraulis japonicus, (27.2%). 11-15,.,,,,. 21-30,, ( ) - -,. 4.6 2.3% ( 3.3%)
(53.8%)
- (13.6%), (8.2%), (6.8%), (6.3%). (,, ) 9%. 11-15, Ц,. 21-35. 2.8%, 2.4-3.5%
- (40%), (34.8%) (15.9%). (,, ),, -, 7.1% 6-10 3.8% 26-30 ( 3.1% )
(42.6%), (25.9%) (23.3%),,.. 21-30,, (31-40 ) Ц, 40. 2.4%, 2.7 1.1 %
- (47.7%), (23.5%) (26.4%). (11-20 ). 21-40, -,,. 41-50 (,, ), -, ( 50 ) -. - 6.8 2.2% ( 3.6%)
(62.2%), -, (32.2%). 11-20. 21-30, (31-50 ),. 50, (,, ). 2.5%, 4.4 2.3%
- (92.5%), (7.2%). 11-20, (,.). 30 -, 41-50. 1.6 3.2% ( 2.5%)
(48.7%),, (35.8%), -,. 11-30.,, (51-60 ). 3.6 1.3%, 2.5%
16-40 - (47.4%), (18.2%), (14.8%) (13.1%).. 30 ( ). 1 0.4% ( 0.4%)
(80.9%). (10%) (4.8%),., 11-15, (36-40 ) -. 1.8 5%, 2.8%
21-50 (98.3%),
,,. (1.7-2%) 1.9%
11-35, (47.8%) (33%). 11-15, ( ). 1.9 4.1 %, 2.8%
(33.7%) (19.6%), - (14%), (11.5%), (7.1%), (7%) (6.5%). 21-30, (31-50 ). 50,. 1-2.1% ( 1.7%)
(75.1%). (17.2%) (5.9%). 11-15, (16-35 ). 1.7%, 3.3 1.3%
(55.8%) (26.1%), - (8.3%), (4.4%) (3.7%). 16Ц20. 21Ц25,,. 26-40, Bivalvia Ц. 1.4 4.2% ( 3.4%)
16-35 (72.5%)
(11.9%), (3.4%), (4.5%) (3%).,, -, Bivalvia,. (3.4-4.2%) 3.9%
16-35 (46.1%) (38%), - (7.5%) (5.2%),. 20,.. 3.3%, 1.9-6.1%
- (40.1%) (38.2%). (5.8%), (4.3%) (4.4%), Ц. 16-20, 21-30, 30. : -,. 1.8 5.4% ( 2.6%)
(64.5%) (31.4%),. 5.3% 11-15 1.6% 35 2.5%
(28.4%) (28.9%), - (16.3%), (14.5%) (6.9%). 11-25 ( ). 26-30, ( ). 30 (, Arctoscopus japonicus.),. 1.4-3.2% ( 2.2%)
(29.9%) (26.8%), - (16%), (3,9%), (7.6%) (6.2%). (11-15 ),. 20. 21-30, (31-35 ). 1.5%, 0.9-1.8 %
4.. 4.1. Ц, (, 1988).,. 3 -, (. 2)
2 :
(,,,,, ) - (, ).,. II, - II-III (
2)
. 2.. : 1 Ц, 2 Ц, 3 - л 9 (,,,,,,, ),. III, - II-III
12, ( - ) (,,,,,,, - ). II-III,, - III-IV, - III
( ). 4.37 4. 3.56 - 4. 3.35 4. 3.94 4. 4.51 3. 3.58 4. 4.12 3. 3.69 3. 4.37 3. 3.61 3. 3.37 3. 4.03 3. 3.88 3. 3.84 3. 3.86 3. - 4.25 3. 3.14 4.58. II-III (34.4%)
(7.2%), (5.3%), (4.5%) (4.3%). - (0.9%) (0.4%). (1.5%) (1.7%), (1.7%) (1%). 2-4%
,, 35.9. (45%). ( 5.1%), (4.6%) (3.1%)
, Ц,.,,. ( )
2-2.2, 1.9, ( 30.3. ). 2.2-2.9%.,, ( - ),.,, - ( ).,
7. (1.9. ), (76.6%). : 2%, - 1.4%. ( ),., 2
.., 3 140.8
, 40.4%, 27% -, 21.3% - 10.5% - (. 3)
. 3. ( ), ( ) ( ). 2 - 76.9.. (40.5%) (27.8%), (18.4%) (13.3%). - 3.2.. ( 43.2 47.9%); 3.9% 5%
. (., 2005;., 2008).,
, (,, 1997;, 2001) Ц. (,, 1941), - 3 (,, 2012),, (, 2010)
, (. 3). II-III ( I-II ),, III III-IV ( II II-III ),., ( ), - (,,, )
(%) ( ). II 29.9 13.6 - 20.II-III 53.9 63.3 65.8 58.III 15.8 2.7 27.4 14.III-IV 0.4 20.4 6.8 7. 2.75 2.85 2.91 2.. 220.9
, ( 32%), - (18.3%) (17%). ( 46.3 31.6%)
(13%) (10.9%), - (17.5%) (14.4%), - (7.7%). 12.3%, - 5.3%, - 2.1% - 2%. -, 42.5%. - (9.6%) (8.2%)
4.2. (, 2006).,.,.. Ц.. (, 2002).,,,,
... Ц.. - ( ).. (1952),.. 4,.. (46%) (47.6%). ( 5.8% 0.6% )
(%) ( ). 50.1 40.7 43. ( 33%) 43 54.1 46. (34-66%) 6.4 4.7 7. ( 66%) 0.5 0.5 2. 4656 3403 4 -, ( 25, 31-35, 30 50, - 21-40 ), - ( 21-30 21- ), ( 11-25 ) ( - 11-20 21-30 )
, ( 26-30, 21-60 11-15, 26-35 21-30, 6-15, 16-25 ), - - ( 31-40, - 30 31-40 60 ) ( 11- 21-30 41-50 ). (26-30 31-35 )
, ( 16-25 21- ), ( 11-30 11-20 ), ( 41-50, 51-60, 26-30 51-60 ) Argis lar lar ( 41-60 31-40 )
, (72%) (27.9%) (. 5). ( 0.09 0.01% )
16- 11-20. 21-25 60 31-40, - 31-40 -
(%). 72.2 71.2 76. ( 1000.) 27.7 28.7 23. (1000-5000.) 0.1 0.09 - ( 5000.) - 0.01 - 9312 6806 9 6-10,. 11-30 41-50, 31-40, 35 16-20, - 31-40.,.,,,
,,., (, 1981)., -, -
. (, 2006),, ( )
1. 32. 11, 9 - 12 Ц. 3.14 4.58. (34.4%), (58.5%) II-III
2. (7.2%), (5.3%), (4.5%) (4.3%), - - (0.9%) (0.4%). 2-4%
3. ( 50 ).,,
4., ( ) (,,, )
5.. 220.., 32%, 18.3% - 17% -. 12.3% - (5.3%), (2.1%) (2%). ( 46.3 31.6%)
6. -, 42.5%. - (9.6%) (8.2%)
7. - (72%) (27.9%).,,
, :
1... ( ) //.. - 1998. Ц. 123. Ц. 185-190
2... Pleurogrammus azonus //.. - 2000. Ц. 127. Ц. 203-208
3...,.. Hexagrammos. ( ) //.. - 2002. Ц. 131. Ц
164-178
4...,.. Myoxocephalus stelleri M. brandti (Cottidae) //.. - 2002. - 42, 4. Ц. 536-542
5...,.. Myoxocephalus (Cottidae). //.. - 2004. Ц. 138. Ц. 120-153
6...,..,.. Bothrocara hollandi (Zoarcidae) ( ) //.. - 2005. Ц. 45, 1. Ц. 81-85
7... //.. - 2005. Ц. 142. Ц. 246-269
8...,..,..,.. Stichaeus grigorjewi (Stichaeidae) //.. - 2006. Ц. 46, 4. Ц. 512-518
9...,..,.. Bathyraja parmifera (Rajidae, Rajiformes) - //.. - 2007. Ц. 47, 3. Ц. 311-319
10...,..,..,.. - Hemitripterus villosus (Pallas, 1814) (Hemitripteridae) - //.. - 2010. Ц. 50, 5. Ц
665-674
11...,..,... Alcichthys elongatus - //.. - 2011. Ц. 51, 2. Ц. 195-204
12...,..,.. //.. - 2011. Ц. 12, 4 (48). Ц. 813-821
13...,..,... Eleginus gracilis (Tilesius, 1810). ( ) //.. - 2011. Ц. 12, 4 (48). Ц. 678-689
, :
14... ( ) //.. - :, 1996. Ц. 121-122
15. Pushchina O.I. The size composition of daily ration of the atca mackerel Pleurogrammus azonus in Primorye coastal waters (Japan sea) // Ninth. Ann. Meet. PICES. - Hakodate, 2000. - P. 120
16...,.. Hexagrammos ( ) //.... - :
, 2001. Ц. 194-196
17...,.. - Myoxocephalus jaok ( ) //.... - :, 2001. Ц. 196-197
18...,..,.. Gadus macrocephalus - - //.,... -. - :
-, 2008. Ц. 127-128
19...,... //. -. :
, 2010.. 1.. 91-95
.-.. 1,0 60 84/ 100. 16
_______________________________________________________________ л - ,.,.,
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